2012
In 2012, bartender Desmond Miles is abducted by Abstergo Industries and forced to use the Animus. Abstergo intends to use Desmond, who is descended from Altaïr, to locate the Apples of Eden and other Isu artefacts. Thankfully for Desmond there’s an Assassin Brotherhood mole working at Abstergo, Lucy Stillman, and she helps him escape from the facility.
Desmond is brought to a Brotherhood safehouse where he works with Lucy and her teammates, Shaun Hastings and Rebecca Crane, to explore the memories of another one of his ancestors: Ezio. Desmond learns of the impending apocalypse via Ezio’s encounter with Minerva beneath the Vatican, kickstarting the Assassins’ quest to prevent the end of the world.
By exploring more of Ezio’s memories, Desmond is able to find the Apple of Eden that was hidden beneath the Roman Colosseum. Unfortunately, the Apple projects a vision of Juno, a villainous Isu who possesses Desmond and forces him to kill Lucy. The traumatic ordeal puts Desmond into a coma, but he’s still able to explore Ezio’s final memories despite such a medical emergency. Through them he observes Ezio’s discovery of Altaïr’s library and Jupiter’s message revealing the existence of the Grand Temple.
Awakening from his coma, Desmond travels with Shaun and Rebecca to the Isu Grand Temple in New York where they meet with William Miles, Desmond’s father and the leader of the modern Assassin Brotherhood. By reliving the memories of Connor Kenway, Desmond is able to locate the key to the Temple. Inside, the group discover that the Isu technology can prevent the apocalypse but at the cost of killing Desmond and unleashing the imprisoned Juno, who plans to control humanity. Desmond decides to sacrifice himself, believing humanity will be able to defeat Juno.
VIA: ign.com